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Author Rubenstein, D. I.; Hack, M. A. doi  openurl
  Title Horse signals: The sounds and scents of fury Type Journal Article
  Year 1992 Publication Evolutionary Ecology Abbreviated Journal (up) Evol. Ecol.  
  Volume 6 Issue 3 Pages 254-260  
  Keywords ommunication – combat – fighting ability – individual identity – signals – information – assessment – displays  
  Abstract During contests animals typically exchange information about fighting ability. Among feral horses these signals involve olfactory or acoustical elements and each type can effectively terminate contests before physical contact becomes necessary. Dung transplant experiments show that for stallions, irrespective of rank, olfactory signals such as dung sniffing encode information about familiarity suggesting that such signals can be used as signatures. As such they can provide indirect information about fighting ability as long as opponents associate identity with past performance. Play-back experiments, however, show that vocalizations, such as squeals, directly provide information about status regardless of stallion familiarity. Sonographs reveal that squeals of dominants are longer than those of subordinates and that only those of dominants have at their onset high-frequency components.  
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  Area Expedition Conference  
  Notes Approved no  
  Call Number refbase @ user @ Serial 506  
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Author Rumbaugh, D.M.; Riesen, A.H.; Wright, S.C. openurl 
  Title Creative responsiveness to objects: a report of a pilot study with young apes Type Journal Article
  Year 1972 Publication Folia Primatologica; International Journal of Primatology Abbreviated Journal (up) Folia Primatol (Basel)  
  Volume 17 Issue 5 Pages 397-403  
  Keywords Animals; *Creativeness; *Hominidae; Pan troglodytes; Play and Playthings; *Problem Solving  
  Abstract  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0015-5713 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:5082622 Approved no  
  Call Number Equine Behaviour @ team @ Serial 4183  
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Author Flack, J.C.; Jeannotte, L.A.; de Waal, F.B.M. doi  openurl
  Title Play signaling and the perception of social rules by juvenile chimpanzees (Pan troglodytes) Type Journal Article
  Year 2004 Publication Journal of comparative psychology (Washington, D.C. : 1983) Abbreviated Journal (up) J Comp Psychol  
  Volume 118 Issue 2 Pages 149-159  
  Keywords Age Factors; Animals; Behavior, Animal; Female; Male; Pan troglodytes; *Play and Playthings; Recognition (Psychology); *Signal Detection (Psychology); *Social Perception  
  Abstract Prescriptive social rules are enforced statistical regularities. The authors investigated whether juvenile chimpanzees (Pan troglodytes) recognize and use enforced statistical regularities to guide dyadic play behavior. They hypothesized (a) that proximity of adults, especially mothers of younger play partners, to play bouts will increase the play signaling of older partners and (b) that when juvenile-juvenile play bouts occur in proximity to adults, older partners will play at a lower intensity than when no adults are present. They found that older and younger partners increase their play signaling in the presence of the mothers of younger partners, particularly as the intensity of play bouts increases. In contrast to their hypothesis, older partners played more roughly when the mothers of younger partners were in proximity. These results suggest that juvenile chimpanzees increase play signaling to prevent termination of the play bouts by mothers of younger partners.  
  Address Santa Fe Institute, NM 97501, USA. jflack@santafe.edu  
  Corporate Author Thesis  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0735-7036 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:15250802 Approved no  
  Call Number refbase @ user @ Serial 172  
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Author Hausberger, M.; Fureix, C.; Bourjade, M.; Wessel-Robert, S.; Richard-Yris, M.-A. url  doi
openurl 
  Title On the significance of adult play: what does social play tell us about adult horse welfare? Type Journal Article
  Year 2012 Publication Abbreviated Journal (up) Naturwissenschaften  
  Volume 99 Issue 4 Pages 291-302  
  Keywords Social play; Stress indicators; Animal welfare; Domestic horse  
  Abstract Play remains a mystery and adult play even more so. More typical of young stages in healthy individuals, it occurs rarely at adult stages but then more often in captive/domestic animals, which can imply spatial, social and/or feeding deprivations or restrictions that are challenging to welfare, than in animals living in natural conditions. Here, we tested the hypothesis that adult play may reflect altered welfare states and chronic stress in horses, in which, as in several species, play rarely occurs at adult stages in natural conditions. We observed the behaviour (in particular, social play) of riding school horses during occasional outings in a paddock and measured several stress indicators when these horses were in their individual home boxes. Our results revealed that (1) the number of horses and rates of adult play appeared very high compared to field report data and (2) most stress indicators measured differed between ‘players’ and ‘non-players’, revealing that most ‘playful’ animals were suffering from more chronic stress than ‘non-playful’ horses. Frequency of play behaviour correlated with a score of chronic stress. This first discovery of a relationship between adult play and altered welfare opens new lines of research that certainly deserves comparative studies in a variety of species.  
  Address  
  Corporate Author Thesis  
  Publisher Springer-Verlag Place of Publication Editor  
  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0028-1042 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5649  
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Author Hausberger, M.; Fureix, C. pdf  openurl
  Title On the significance of puzzling behaviours: what do yawning and adult play tell us about horse (Equus caballus) welfare? Type Conference Article
  Year 2012 Publication Proceedings of the 2. International Equine Science Meeting Abbreviated Journal (up) Proc. 2. Int. Equine. Sci. Mtg  
  Volume in press Issue Pages  
  Keywords welfare indicators, yawning, adult play, altered welfare, horse  
  Abstract Some behaviours remain a mystery and their determinants are still uncertain. Here we propose a critical interpretation of two puzzling behaviours, i.e. yawning and adult play, recorded in large samples of domestic horses kept in sub-optimal conditions (e.g. time-restricted feeding practices, social isolation). Yawning is involved in behavioural state changes, especially in quiet contexts of motor relaxation, but seems also to be triggered by stress and emotional contexts. Thus, we investigated specific correlates of yawning and stereotypic behaviours in two large samples of working horses (n1 = 87, n2 = 59). A clear co-occurrences of yawning and stereotypic behaviours appeared: stereotypic horses yawned more than the non-stereotypic horses (Fisher test, p < 0.02), yawning increased at the same time periods as stereotypic behaviours did (Wilcoxon tests, p < 0.05 to 0.001) and yawning frequency was even sometimes positively correlated with stereotypic behaviour frequencies (Spearman correlations, p < 0.005). Play, more typical of young stages in healthy individuals, occurs rarely at adult stages but then more often in captive/domestic animals (which can be challenging to welfare) than in animals living in natural conditions. Thus, we tested the hypothesis that adult play may reflect altered welfare states in horses, where, as in several species, play rarely occurs at adult stages in natural conditions. We observed the behaviour (in particular social play) of 29 adult riding school horses during occasional outings in a paddock and measured several stress indicators (health-related, behavioural and physiological) when these horses were in their home individual boxes. Results revealed that the number of horses and rates of adult play appeared very high compared to field report data and that most stress indicators measured differed between “players” and “non-players”, revealing that most “playful” animals were more prone to suffer from chronic stress contrarily to “non-playful” horses (Fisher, Chi-square, Mann-Whitney and Spearman correlation tests, p < 0.05 to 0.001). Altogether, these results show that yawning and adult play can both reflect altered welfare states in horses, highlighting that these puzzling behaviours should not be considered, as currently thought, as reliable indicators of good welfare states and / or expression of positive emotions.  
  Address  
  Corporate Author Hausberger, M. Thesis  
  Publisher Xenophon Publishing Place of Publication Wald Editor Krueger, K.  
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  ISSN 978-3-9808134-26 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5516  
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Author Proops, L.; McComb, K.; Reby, D. url  doi
openurl 
  Title Cross-modal individual recognition in domestic horses (Equus caballus) Type Journal Article
  Year 2009 Publication Proceedings of the National Academy of Sciences of the United States of America Abbreviated Journal (up) Proc. Natl. Acad. Sci. U.S.A.  
  Volume 106 Issue 3 Pages 947-951  
  Keywords animal cognition vocal communication social behavior playback experiment expectancy violation  
  Abstract Individual recognition is considered a complex process and, although it is believed to be widespread across animal taxa, the cognitive mechanisms underlying this ability are poorly understood. An essential feature of individual recognition in humans is that it is cross-modal, allowing the matching of current sensory cues to identity with stored information about that specific individual from other modalities. Here, we use a cross-modal expectancy violation paradigm to provide a clear and systematic demonstration of cross-modal individual recognition in a nonhuman animal: the domestic horse. Subjects watched a herd member being led past them before the individual went of view, and a call from that or a different associate was played from a loudspeaker positioned close to the point of disappearance. When horses were shown one associate and then the call of a different associate was played, they responded more quickly and looked significantly longer in the direction of the call than when the call matched the herd member just seen, an indication that the incongruent combination violated their expectations. Thus, horses appear to possess a cross-modal representation of known individuals containing unique auditory and visual/olfactory information. Our paradigm could provide a powerful way to study individual recognition across a wide range of species.  
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  Notes 10.1073/pnas.0809127105 Approved no  
  Call Number Equine Behaviour @ team @ Serial 4689  
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Author Kawamura, S. openurl 
  Title Aggression as studied in troops of Japanese monkeys Type Journal Article
  Year 1967 Publication UCLA Forum in Medical Sciences Abbreviated Journal (up) UCLA Forum Med Sci  
  Volume 7 Issue Pages 195-223  
  Keywords *Aggression; Animals; Behavior, Animal; Defense Mechanisms; Female; *Haplorhini; Homosexuality; Humans; Japan; Leadership; Male; Play and Playthings; Sexual Behavior, Animal; Social Behavior; Socialization  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0082-7134 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:4972333 Approved no  
  Call Number Serial 2056  
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Author Beaver, B.V. openurl 
  Title Aggressive behavior problems Type Journal Article
  Year 1986 Publication The Veterinary clinics of North America. Equine practice Abbreviated Journal (up) Vet Clin North Am Equine Pract  
  Volume 2 Issue 3 Pages 635-644  
  Keywords Affect; Aggression/*psychology; Animals; *Behavior, Animal; Dominance-Subordination; Fear; *Horses; Play and Playthings; Sexual Behavior, Animal; Social Environment  
  Abstract Accurate diagnosis of the cause of aggression in horses is essential to determining the appropriate course of action. The affective forms of aggression include fear-induced, pain-induced, intermale, dominance, protective, maternal, learned, and redirected aggressions. Non-affective aggression includes play and sex-related forms. Irritable aggression and hypertestosteronism in mares are medical problems, whereas genetic factors, brain dysfunction, and self-mutilation are also concerns.  
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  Series Volume Series Issue Edition  
  ISSN 0749-0739 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:3492250 Approved no  
  Call Number refbase @ user @ Serial 674  
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