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	Author	Broom, M.
	Title	A unified model of dominance hierarchy formation and maintenance			Type	Journal Article
	Year	2002	Publication	Journal of theoretical biology	Abbreviated Journal	J. Theor. Biol.
	Volume	219	Issue	1	Pages	63-72
	Keywords	Animals; Behavior, Animal; Feeding Behavior; Models, Psychological; *Social Dominance; Social Environment
	Abstract	In many different species it is common for animals to spend large portions of their lives in groups. Such groups need to divide available resources amongst the individuals they contain and this is often achieved by means of a dominance hierarchy. Sometimes hierarchies are stable over a long period of time and new individuals slot into pre-determined positions, but there are many situations where this is not so and a hierarchy is formed out of a group of individuals meeting for the first time. There are several different models both of the formation of such dominance hierarchies and of already existing hierarchies. These models often treat the two phases as entirely separate, whereas in reality, if there is a genuine formation phase to the hierarchy, behaviour in this phase will be governed by the rewards available, which in turn depends upon how the hierarchy operates once it has been formed. This paper describes a method of unifying models of these two distinct phases, assuming that the hierarchy formed is stable. In particular a framework is introduced which allows a variety of different models of each of the two parts to be used in conjunction with each other, thus enabling a wide range of situations to be modelled. Some examples are given to show how this works in practice.
	Address	Centre for Statistics and Stochastic Modelling, School of Mathematical Sciences, University of Sussex, Falmer, Brighton, BN1 9QH, U.K. m.broom@sussex.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5193	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12392975			Approved	no
	Call Number	refbase @ user @			Serial	439
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	Author	Viscido, S.V.; Miller, M.; Wethey, D.S.
	Title	The dilemma of the selfish herd: the search for a realistic movement rule			Type	Journal Article
	Year	2002	Publication	Journal of theoretical biology	Abbreviated Journal	J. Theor. Biol.
	Volume	217	Issue	2	Pages	183-194
	Keywords	Animals; Behavior, Animal; Mass Behavior; Models, Biological; *Motor Activity; Predatory Behavior
	Abstract	The selfish herd hypothesis predicts that aggregations form because individuals move toward one another to minimize their own predation risk. The “dilemma of the selfish herd” is that movement rules that are easy for individuals to follow, fail to produce true aggregations, while rules that produce aggregations require individual behavior so complex that one may doubt most animals can follow them. If natural selection at the individual level is responsible for herding behavior, a solution to the dilemma must exist. Using computer simulations, we examined four different movement rules. Relative predation risk was different for all four movement rules (p<0.05). We defined three criteria for measuring the quality of a movement rule. A good movement rule should (a) be statistically likely to benefit an individual that follows it, (b) be something we can imagine most animals are capable of following, and (c) result in a centrally compact flock. The local crowded horizon rule, which allowed individuals to take the positions of many flock-mates into account, but decreased the influence of flock-mates with distance, best satisfied these criteria. The local crowded horizon rule was very sensitive to the animal's perceptive ability. Therefore, the animal's ability to detect its neighbors is an important factor in the dynamics of group formation.
	Address	Department of Biological Sciences, University of South Carolina, Columbia, SC, 29208, USA. viscido@u.washington.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5193	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12202112			Approved	no
	Call Number	refbase @ user @			Serial	554
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	Author	Viscido, S.V.; Miller, M.; Wethey, D.S.
	Title	The response of a selfish herd to an attack from outside the group perimeter			Type	Journal Article
	Year	2001	Publication	Journal of theoretical biology	Abbreviated Journal	J. Theor. Biol.
	Volume	208	Issue	3	Pages	315-328
	Keywords	Animals; Behavior, Animal; Computer Simulation; Models, Biological; *Movement; Predatory Behavior
	Abstract	According to the selfish herd hypothesis, animals can decrease predation risk by moving toward one another if the predator can appear anywhere and will attack the nearest target. Previous studies have shown that aggregations can form using simple movement rules designed to decrease each animal's Domain of Danger. However, if the predator attacks from outside the group's perimeter, these simple movement rules might not lead to aggregation. To test whether simple selfish movement rules would decrease predation risk for those situations when the predator attacks from outside the flock perimeter, we constructed a computer model that allowed flocks of 75 simulated fiddler crabs to react to one another, and to a predator attacking from 7 m away. We attacked simulated crab flocks with predators of different sizes and attack speeds, and computed relative predation risk after 120 time steps. Final trajectories showed flight toward the center of the flock, but curving away from the predator. Path curvature depended on the predator's size and approach speed. The average crab experienced a greater decrease in predation risk when the predator was small or slow moving. Regardless of the predator's size and speed, however, predation risk always decreased as long as crabs took their flock-mates into account. We conclude that, even when flight away from an external predator occurs, the selfish avoidance of danger can lead to aggregation.
	Address	Department of Biological Sciences, University of South Carolina, Columbia, SC 29208, U.S.A. viscido@u.washington.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5193	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11207093			Approved	no
	Call Number	refbase @ user @			Serial	555
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	Author	Suzuki, Y.; Toquenaga, Y.
	Title	Effects of information and group structure on evolution of altruism: analysis of two-score model by covariance and contextual analyses			Type	Journal Article
	Year	2005	Publication	Journal of theoretical biology	Abbreviated Journal	J. Theor. Biol.
	Volume	232	Issue	2	Pages	191-201
	Keywords	Altruism; Analysis of Variance; Communication; Cooperative Behavior; Evolution; Game Theory; Group Structure; Humans; Models, Genetic; Models, Psychological; Selection (Genetics); Trust
	Abstract	An altruistic individual has to gamble on cooperation to a stranger because it does not know whether the stranger is trustworthy before direct interaction. Nowak and Sigmund (Nature 393 (1998a) 573; J. Theor. Biol. 194 (1998b) 561) presented a new theoretical framework of indirect reciprocal altruism by image scoring game where all individuals are informed about a partner's behavior from its image score without direct interaction. Interestingly, in a simplified version of the image scoring game, the evolutionarily stability condition for altruism became a similar form of Hamilton's rule, i.e. inequality that the probability of getting correct information is more than the ratio of cost to benefit. Since the Hamilton's rule was derived by evolutionarily stable analysis, the evolutionary meaning of the probability of getting correct information has not been clearly examined in terms of kin and group selection. In this study, we applied covariance analysis to the two-score model for deriving the Hamilton's rule. We confirmed that the probability of getting correct information was proportional to the bias of altruistic interactions caused by using information about a partner's image score. The Hamilton's rule was dependent on the number of game bouts even though the information reduced the risk of cooperation to selfish one at the first encounter. In addition, we incorporated group structure to the two-score model to examine whether the probability of getting correct information affect selection for altruism by group selection. We calculated a Hamilton's rule of group selection by contextual analysis. Group selection is very effective when either the probability of getting correct information or that of future interaction, or both are low. The two Hamilton's rules derived by covariance and contextual analyses demonstrated the effects of information and group structure on the evolution of altruism. We inferred that information about a partner's behavior and group structure can produce flexible pathways for the evolution of altruism.
	Address	Integrative Environmental Sciences, Graduate School of Life and Environmental Sciences, University of Tsukuba, 1-1-1, Ten-Nou-Dai, Tsukuba, Ibaraki 305-8572, Japan. yukari@pe.ies.life.tsukuba.ac.jp
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5193	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15530489			Approved	no
	Call Number	refbase @ user @			Serial	556
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	Author	Hamilton, W.D.
	Title	Geometry for the selfish herd			Type	Journal Article
	Year	1971	Publication	Journal of theoretical biology	Abbreviated Journal	J. Theor. Biol.
	Volume	31	Issue	2	Pages	295-311
	Keywords	Animals; Anura; Behavior, Animal; Breeding; Communication; Evolution; Fear; Metallurgy; Models, Biological; Probability; Snakes; *Spatial Behavior
	Abstract	This paper presents an antithesis to the view that gregarious behaviour is evolved through benefits to the population or species. Following Galton (1871) and Williams (1964) gregarious behaviour is considered as a form of cover-seeking in which each animal tries to reduce its chance of being caught by a predator. It is easy to see how pruning of marginal individuals can maintain centripetal instincts in already gregarious species; some evidence that marginal pruning actually occurs is summarized. Besides this, simply defined models are used to show that even in non-gregarious species selection is likely to favour individuals who stay close to others. Although not universal or unipotent, cover-seeking is a widespread and important element in animal aggregation, as the literature shows. Neglect of the idea has probably followed from a general disbelief that evolution can be dysgenic for a species. Nevertheless, selection theory provides no support for such disbelief in the case of species with outbreeding or unsubdivided populations. The model for two dimensions involves a complex problem in geometrical probability which has relevance also in metallurgy and communication science. Some empirical data on this, gathered from random number plots, is presented as of possible heuristic value.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5193	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:5104951			Approved	no
	Call Number	refbase @ user @			Serial	771
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	Author	Seyfarth, R.M.
	Title	A model of social grooming among adult female monkeys			Type	Journal Article
	Year	1977	Publication	Journal of Theoretical Biology	Abbreviated Journal	J. Theor. Biol.
	Volume	65	Issue	4	Pages	671-698
	Keywords	Animals; Behavior, Animal; Female; Grooming; Haplorhini/physiology; *Models, Biological; Reproduction; Social Dominance; Time Factors
	Abstract	Grooming networks among adult female monkeys exhibit two similar features across a number of different species. High-ranking animals receive more grooming than others, and the majority of grooming occurs between females of adjacent rank. A theoretical model which duplicates these features is presented, and the properties of the model are used to explain the possible causation and function of female grooming behaviour. The model illustrates how relatively simple principles governing the behaviour of individuals may be used to explain more complex aspects of the social structure of non-human primate groups.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5193	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:406485			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5259
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	Author	Nicol, C.J.
	Title	Development, direction, and damage limitation: social learning in domestic fowl			Type	Journal Article
	Year	2004	Publication	Learning & behavior : a Psychonomic Society publication	Abbreviated Journal	Learn Behav
	Volume	32	Issue	1	Pages	72-81
	Keywords	Adaptation, Psychological; Age Factors; Animals; Behavior, Animal; Chickens; Feeding Behavior; Food Preferences; Imitative Behavior; Imprinting (Psychology); Learning; Maternal Behavior; Reinforcement (Psychology); Social Environment; *Social Facilitation
	Abstract	This review highlights two areas of particular interest in the study of social learning in fowl. First, the role of social learning in the development of feeding and foraging behavior in young chicks and older birds is described. The role of the hen as a demonstrator and possible teacher is considered, and the subsequent social influence of brood mates and other companions on food avoidance and food preference learning is discussed. Second, the way in which work on domestic fowl has contributed to an understanding of the importance of directed social learning is examined. The well-characterized hierarchical social organization of small chicken flocks has been used to design studies which demonstrate that the probability of social transmission is strongly influenced by social relationships between birds. The practical implications of understanding the role of social learning in the spread of injurious behaviors in this economically important species are briefly considered.
	Address	Department of Clinical Veterinary Science, University of Bristol, Langford, Bristol, England. c.j.nicol@bristol.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1543-4494	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15161142			Approved	no
	Call Number	refbase @ user @			Serial	75
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	Author	Zentall, T.R.
	Title	Action imitation in birds			Type	Journal Article
	Year	2004	Publication	Learning & behavior : a Psychonomic Society publication	Abbreviated Journal	Learn Behav
	Volume	32	Issue	1	Pages	15-23
	Keywords	Adaptation, Psychological; Animals; Birds; Imitative Behavior; Imprinting (Psychology); Learning; Motivation; Psychological Theory; Social Environment; *Social Facilitation; Vocalization, Animal
	Abstract	Action imitation, once thought to be a behavior almost exclusively limited to humans and the great apes, surprisingly also has been found in a number of bird species. Because imitation has been viewed by some psychologists as a form of intelligent behavior, there has been interest in how it is distributed among animal species. Although the mechanisms responsible for action imitation are not clear, we are now at least beginning to understand the conditions under which it occurs. In this article, I try to identify and differentiate the various forms of socially influenced behavior (species-typical social reactions, social effects on motivation, social effects on perception, socially influenced learning, and action imitation) and explain why it is important to differentiate imitation from other forms of social influence. I also examine some of the variables that appear to be involved in the occurrence of imitation. Finally, I speculate about why a number of bird species, but few mammal species, appear to imitate.
	Address	Department of Psychology, University of Kentucky, Lexington, Kentucky 40506, USA. zentall@uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1543-4494	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15161137			Approved	no
	Call Number	refbase @ user @			Serial	230
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	Author	Gibson, B.M.; Shettleworth, S.J.
	Title	Competition among spatial cues in a naturalistic food-carrying task			Type	Journal Article
	Year	2003	Publication	Learning & behavior : a Psychonomic Society publication	Abbreviated Journal	Learn Behav
	Volume	31	Issue	2	Pages	143-159
	Keywords	Adaptation, Psychological; Animals; Appetitive Behavior; Association Learning; Attention; Choice Behavior; Cues; Discrimination Learning; Male; Rats; Rats, Long-Evans; Space Perception; *Spatial Behavior
	Abstract	Rats collected nuts from a container in a large arena in four experiments testing how learning about a beacon or cue at a goal interacts with learning about other spatial cues (place learning). Place learning was quick, with little evidence of competition from the beacon (Experiments 1 and 2). Rats trained to approach a beacon regardless of its location were subsequently impaired when the well-learned beacon was removed and other spatial cues identified the location of the goal (Experiment 3). The competition between beacon and place cues reflected learned irrelevance for place cues (Experiment 4). The findings differ from those of some studies of associative interactions between cue and place learning in other paradigms.
	Address	University of Toronto, Toronto, Ontario, Canada
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1543-4494	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12882373			Approved	no
	Call Number	refbase @ user @			Serial	368
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	Author	Fragaszy, D.; Visalberghi, E.
	Title	Socially biased learning in monkeys			Type	Journal Article
	Year	2004	Publication	Learning & behavior : a Psychonomic Society publication	Abbreviated Journal	Learn Behav
	Volume	32	Issue	1	Pages	24-35
	Keywords	Adaptation, Psychological; Animal Communication; Animals; Behavior, Animal; Feeding Behavior/psychology; Food Preferences/psychology; Haplorhini/psychology; Imitative Behavior; Imprinting (Psychology); Learning; Social Environment; Social Facilitation
	Abstract	We review socially biased learning about food and problem solving in monkeys, relying especially on studies with tufted capuchin monkeys (Cebus apella) and callitrichid monkeys. Capuchin monkeys most effectively learn to solve a new problem when they can act jointly with an experienced partner in a socially tolerant setting and when the problem can be solved by direct action on an object or substrate, but they do not learn by imitation. Capuchin monkeys are motivated to eat foods, whether familiar or novel, when they are with others that are eating, regardless of what the others are eating. Thus, social bias in learning about foods is indirect and mediated by facilitation of feeding. In most respects, social biases in learning are similar in capuchins and callitrichids, except that callitrichids provide more specific behavioral cues to others about the availability and palatability of foods. Callitrichids generally are more tolerant toward group members and coordinate their activity in space and time more closely than capuchins do. These characteristics support stronger social biases in learning in callitrichids than in capuchins in some situations. On the other hand, callitrichids' more limited range of manipulative behaviors, greater neophobia, and greater sensitivity to the risk of predation restricts what these monkeys learn in comparison with capuchins. We suggest that socially biased learning is always the collective outcome of interacting physical, social, and individual factors, and that differences across populations and species in social bias in learning reflect variations in all these dimensions. Progress in understanding socially biased learning in nonhuman species will be aided by the development of appropriately detailed models of the richly interconnected processes affecting learning.
	Address	Psychology Department, University of Georgia, Athens, Georgia 30602, USA. doree@uga.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1543-4494	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15161138			Approved	no
	Call Number	refbase @ user @			Serial	828
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