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Author Kirkpatrick, J.F.; Turner, J.W. doi  openurl
  Title Changes in herd stallions among feral horse bands and the absence of forced copulation and induced abortion Type Journal Article
  Year 1991 Publication Behavioral Ecology and Sociobiology Abbreviated Journal Behav. Ecol. Sociobiol.  
  Volume 29 Issue 3 Pages 217-219  
  Keywords  
  Abstract Forced copulation and induced abortion were investigated in a herd of feral horses inhabiting a coastal barrier island. Eight mares were diagnosed pregnant in August and October 1989 by means of urinary and fecal steroid metabolites, prior to documented changes in herd stallions. These mares were observed for harassment and forced copulation by the new stallions and for the presence of foals during the spring and summer of 1990. No incidents of harassment or attempts at forced copulation were witnessed and seven of the eight mares produced foals in 1990. These data indicate that forced copulation and induced abortion are not common events among all feral horse herds and suggest reinvestigation of this hypothesized phenomenon.  
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  Call Number Equine Behaviour @ team @ Serial 2327  
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Author Garott, R.A. doi  openurl
  Title Sex Ratios and Differential Survival of Feral Hors Type Journal Article
  Year 1991 Publication Journal of Animal Ecology Abbreviated Journal J Anim Ecol  
  Volume 60 Issue 3 Pages 929-936  
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  Abstract (1) Sex and age data were collected on 60 111 feral horses (Equus caballus L.) removed from eighty-nine areas in Nevada, Wyoming, and Oregon between 1976 and 1987. (2) Sex ratios of young seldom differed from parity; however, sex ratios of adults were commonly skewed toward females. No evidence of differential capture probability between adult males and females could be detected; therefore, skewed adult sex ratios were attributed to differential survival. (3) Age-specific trends in sex ratios indicated that the proportion of males steadily decreased from near parity in foals, to lows of 0.61-0.77 in the 4-5-year age-classes. The trend then reversed with males becoming predominant (1.08-1.36) in the > 10 years age-class. (4) Population simulations suggest that survival diffentials of 0.05-0.07, favouring females to 4 years of age, and 0.02-0.04 favouring males in older age-classes were required to mimic observed age-specific sex ratio changes. To obtain the high proportion of males in the > 10-years age-class, onset of senescence also had to be earlier for females. (5) Causes for differential survival in the immature age-classes are uncertain, but may relate to behavioural or metabolic differences between the sexes. Differential survival between adult males and females is attributed to differences in the energetic costs of reproduction and disparity in their reproductive life spans.  
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  Call Number Equine Behaviour @ team @ Serial 2294  
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Author Lucas, Z.; Raeside, J.I.; Betteridge, K.J. openurl 
  Title Non-invasive assessment of the incidences of pregnancy and pregnancy loss in the feral horses of Sable Island Type Journal Article
  Year 1991 Publication Journal of Reproduction and Fertility. Supplement Abbreviated Journal J Reprod Fertil Suppl  
  Volume 44 Issue Pages 479-488  
  Keywords Abortion, Veterinary/*epidemiology; Animals; Animals, Wild/*physiology; Atlantic Islands; Estrogens/analysis; Feces/chemistry; Female; Fertilization; Gestational Age; Horse Diseases/*epidemiology; Horses; Incidence; Pregnancy; *Pregnancy, Animal  
  Abstract Field observations of 400 totally unmanaged feral horses on Sable Island, Nova Scotia, were complemented by oestrogen determinations in faecal samples from 154 identified females over a 4-year period (454 mare-years). Of mares that were sampled throughout the year and subsequently produced foals, 92.1% exhibited elevated faecal oestrogens between 15 October and 30 March. The results confirm that faecal oestrogens are a useful indicator of pregnancy after approximately 120 days gestation. Distribution of foaling resembled that seen in other feral populations, with 95% of births occurring from April through July. The foaling rate for mares aged 3 years or older was 62.0%, with 50.7% of mares foaling in 3 or 4 years. Foaling rates were low (4.1%) in mares bred as yearlings and rose with age to 70.8% in those bred as 4-year-olds. Fetal loss after Day 120 was deduced from faecal oestrogens to be 26.0% overall, with marked variation from year to year (9.6-37.3%) and with age (70.0% in those bred as yearlings, decreasing to 5.6% in those bred as 4-year-olds). Of 58 mares aged 2 years or older that were sampled every year, about half (49.6%) the barren years were attributable to fetal loss after 120 days gestation. All mares conceived in at least 2 of the 4 years, suggesting that pregnancy loss, even after Day 120, is as important as failure to conceive in causing barren years.  
  Address Department of Biomedical Sciences, Ontario Veterinary College, University of Guelph, Canada  
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  ISSN 0449-3087 ISBN Medium  
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  Notes PMID:1795292 Approved no  
  Call Number Equine Behaviour @ team @ Serial 2247  
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Author Ben-Shahar, R doi  openurl
  Title Selectivity in large generalist herbivores: feeding patterns of African ungulates in a semi-arid habitat Type Journal Article
  Year 1991 Publication African Journal of Ecology Abbreviated Journal Afr. J. Ecol.  
  Volume 29 Issue 4 Pages 302-315  
  Keywords diet; forage production; grazing; wildebeest; zebra  
  Abstract Feeding habits of free-ranging wildebeest and zebra were monitored in a semi-arid nature reserve, bordering the southwestern part of Kruger National Park, South Africa. The purpose of study was to distinguish and define the feeding niches of two roughage grazers that occur in similar habitat types. The monthly compositions of diets were evaluated by direct observations of feeding bouts over a period of two years when rainfall patterns were average and animal populations were stable. Other analyses evaluated the standing biomass of grass species in the reserve during the wet summer and dry winter seasons.

A considerable overlap of grass species composition was found in the diets of wildebeest and zebra. Ordination of bi-monthly records of the diet composition showed greater variations in scores of grasses in zebra diet in comparison to wildebeest. Seasonal patterns were more apparent in the wildebeest diet. Preference ranking of grass species indicated that zebra diet remained constant in winter and summer. Wildebeest diet however, alternated with seasons, showing high preferences during the winter months for grass species which were rejected during summer.

The combined assessment of results from three separate statistical methods analysing temporal patterns and preferences in diet composition revealed contradictory trends. The solution, however, relied on the initial assumptions posed. Hence, wildebeest and zebra are essentially generalist feeders which show a limited amount of preference in their choice of diet.
 
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  Call Number Equine Behaviour @ team @ Serial 2226  
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Author Boyd, L.E. url  doi
openurl 
  Title The behaviour of Przewalski's horses and its importance to their management Type Journal Article
  Year 1991 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 29 Issue 1-4 Pages 301-318  
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  Abstract Przewalski's horses (Equus przewalskii) are believed to be extinct in the wild; the current known population of 797 animals exists wholly in zoos. The Species Survival Commission of the International Union for Conservation of Nature and Natural Resources is proposing to reintroduce this endangered species into its former Mongogian habitat within the next decade. Knowledge of the behavior of harem-forming equids in general and of Przewalski's horses in particular, is of great importance to the captive propagation and eventual reintroduction of this species. Horses are rarely solitary by nature. Solitary captive animals are prone to pacing. Juvenile male feral horses (Equus caballus) form bachelor herds upon dispersal from their natal band. Zoos can set up bachelor herds as a way of managing surplus males. The older, more dominant feral horse bachelors are the first to acquire mares. Bachelors do not generally obtain females until they are 4 or 5 years of age. The first females acquired are usually 1- and 2-year-old fillies dispersing from their natal band. Because of the age differential, the stallions are generally dominant to their mares. Behavioral impotence may result if captive stallions are given a harem at too young an age, especially if the harem contains older, more dominant, females. Typical harem sizes in the wild are 3-5 mares. Captive stallions with too large a harem may become either apathetic or aggressive toward their mares. Wild horses spend 60-70% of their time foraging. Captive animals may quickly consume their limited amounts of food and develop vices out of boredom. Provision of hay ad libitum reduces the amount of pacing seen in captive animals, and virtually eliminates coprophagy.  
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  Call Number Equine Behaviour @ team @ Serial 2214  
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Author Dugatkin, L.A. doi  openurl
  Title Dynamics of the TIT FOR TAT strategy during predator inspection in the guppy (Poecilia reticulata) Type Journal Article
  Year 1991 Publication Behavioral Ecology and Sociobiology Abbreviated Journal Behav. Ecol. Sociobiol.  
  Volume 29 Issue 2 Pages 127-132  
  Keywords  
  Abstract One well-known solution to the iterated Prisoner's Dilemma is the TIT FOR TAT strategy. This strategy has three “characteristics” associated with it. TIT FOR TAT is nice (cooperates on the first move of a game), retaliatory (plays defect against an individual that defected on the prior move), and forgiving (cooperates with an individual which has defected in the past but cooperates in the present). Predator inspection behavior in guppies (Poecilia reticulata) was examined in order to determine whether guppies displayed these three characteristics. Results indicate that while it can be quite difficult to translate the abstract concepts of niceness, retaliation, and forgiveness into measurable behaviors, the data support the hypothesis that guppies display the three characteristics associated with the TIT FOR TAT strategy.  
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  Call Number Equine Behaviour @ team @ Serial 2178  
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Author Dugatkin, L.; Alfieri, M. doi  openurl
  Title Tit-For-Tat in guppies (Poecilia reticulata): the relative nature of cooperation and defection during predator inspection Type Journal Article
  Year 1991 Publication Evolutionary Ecology Abbreviated Journal Evol. Ecol.  
  Volume 5 Issue 3 Pages 300-309  
  Keywords Game theory – Tit-For-Tat – predator inspection – guppy  
  Abstract Summary The introduction of game-theoretical thinking into evolutionary biology has laid the groundwork for a heuristic view of animal behaviour in which individuals employ “strategies” – rules that instruct them how to behave in a given circumstance to maximize relative fitness. Axelrod and Hamilton (1981) found that a strategy called Tit-For-Tat (TFT) is one robust cooperative solution to the iterated Prisoner's Dilemma game. There exists, however, little empirical evidence that animals employ TFT. Predator inspection in fish provides one ecological context in which to examine the use of the TFT strategy.  
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  Call Number Equine Behaviour @ team @ Serial 2177  
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Author Beauchamp, G.; Kacelnik, A. url  doi
openurl 
  Title Effects of the knowledge of partners on learning rates in zebra finches Taeniopygia guttata Type Journal Article
  Year 1991 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.  
  Volume 41 Issue 2 Pages 247-253  
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  Abstract Many interpretations of the adaptive value of group living involve tranfer of knowledge. However, according to learning theory, being in a pair with a knowledgeable partner can have paradoxical consequences. Obtaining food by following a skilled companion may reduce the ability of naive individuals to learn about clues that signal the occurrence of food. This study examined the relation between learning and following in paris of zebra finches. Knowledgeable partners were trained to obtain food from a computer-controlled dispenser by using the information provided by a signal. For non-knowledgeable partners, the signal was irrelevant and could not be used to predict foraging opportunities. The rate of learning about the signal by naive birds that shared the experience of either knowledgeable or nonknowledgeable tutors was then examined. Naive birds learned more slowly as a result of being in a pair with a knowledgeable than a non-knowledgeable partner. Well-informed mates acted as a reliable cue to predict foraging opportunities, and thus overshadowed the independent signal. The knowledge of a partner influences learning rates in naive individuals, but in the opposite direction to that predicted by earlier accounts of learning in social contexts.  
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  Notes Approved no  
  Call Number Serial 2115  
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Author Walther, F.R. url  openurl
  Title On herding behavior Type Journal Article
  Year 1991 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 29 Issue 1-4 Pages 5-13  
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  Abstract Herding behavior in ungulates is executed mainly by males. There are several forms of herding: guarding a single estrous female; rounding up a bunch of females during the rutting season; territorial herding by which a male keeps females inside his territory; herding of a moving, permanent, harem group; social herding in which group members of both sexes are herded by one dominant male. When put into this sequence, a phenotypical trend is illustrated, leading from an intimate connection of herding with mating behavior toward an increasing independence from sexual behavior and culminating in a complete socialization of herding. Aspects and problems of herding behavior are the recognition of, and the animal's respect for, partners in gregarious species; the animal “taking offense” at activities of others which deviate from its own activity; the active coordination and synchronization of group activities; the use of, and the effects of, threat, dominance and courtship displays in herding; the possibility of substitution among expressive displays in relation to partners of different sex; the communicative function of the animal's orientation relative to the partner; social hierarchy and leadership in a group; the possibility of interspecific herding, particularly in man-animal relationships, which is closely linked to the process of domestication.  
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  Notes Approved no  
  Call Number Serial 2035  
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Author Kabuga, J.D.; Gari-Kwaku, J.; Annor, S.Y. url  doi
openurl 
  Title Social status and its relationships to maintenance behaviour in a herd of N'dama and West African Shorthorn cattle Type Journal Article
  Year 1991 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 31 Issue 3-4 Pages 169-181  
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  Abstract Social-related behaviour of N'dama and West African Shorthorn (WASH) cattle grazing together was studied over 10 consecutive days. Dominance was not related to age, liveweight or leadership when the animals were led into a weighing scale or into and out of the experimental paddock. Dominance had no influence on the use of shade, on drinking frequency or on grazing time, it was, however, positively associated with time spent ruminating and idling and with the frequency of allogrooming. Forced leadership into a weighing scale was negatively correlated (Spearmans rank correlation (rs=-0.69, P<0.05) with liveweight while voluntary leadership, out of the experimental paddock (rs=0.85, P<0.01) and into the experimental paddock (rs=0.76, P<0.05), was positively correlated with liveweight. Voluntary leadership also positively and significantly (P<0.01) influenced the frequency of visits to the water trough. All measures of leadership were significantly but negatively correlated with frequency of social association (close contact) between cows. N'dama were more aggressive than WASH and had higher dominance values. There was a slight tendency for WASH to associate more with their peers than N'dama with their peers. Social behaviours such as allogrooming were low and rubbing and sniffing absent in both breeds.  
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  Notes Approved no  
  Call Number Serial 2034  
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