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Author Hanggi, E.B.; Ingersoll, J.F. url  doi
openurl 
  Title Long-term memory for categories and concepts in horses (Equus caballus) Type Journal Article
  Year 2009 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume 13 Issue 3 Pages 451-462  
  Keywords Horse – Long-term memory – Concept – Categorization – Discrimination – LCD display – Equine  
  Abstract Three horses (Equus caballus) with a history of performing cognitive tasks including discrimination learning, categorization, and concept use were tested to evaluate their long-term memory (LTM) in three experiments. In addition, use of LCD multi-displays for stimulus presentation was incorporated into cognition testing protocol for the first time with horses. Experiment 1 tested LTM for discrimination learning that originally occurred 6 years earlier. Five sets of stimuli were used and the two horses tested showed no decrement in performance on four of the sets; however, both horses did score below chance on one set. Experiment 2 examined long-term categorization recall 10 years after horses had demonstrated the ability to make stimulus selections based on shared characteristics within a given category. The horse tested for LTM after the decade-long interval immediately and consistently applied the previously learned categorization rule to not only familiar but also novel sets of stimuli. Experiment 3 tested another horse for LTM for a relative size concept. This horse had originally demonstrated concept rule use in order to select stimuli based on their relative size to one another. More than 7 years later and without further training, this horse reliably applied the previously established size concept to both familiar and novel sets of stimuli. These findings are the first reports of long-term categorical and conceptual memory in horses and are consistent with observations of domestic and wild horses, which indicate that behavioral and ecological events may be remembered for long periods of time. These studies also demonstrate the adaptive nature of horses with regard to their ability to generalize over several different testing conditions.  
  Address Equine Research Foundation, P.O. Box 1900, Aptos, CA, 95001, USA, EquiResF@aol.com  
  Corporate Author Thesis  
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  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1435-9456 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:19148689 Approved no  
  Call Number Equine Behaviour @ team @ Serial 4760  
Permanent link to this record
 
 
Author Dubois, C.; Manfredi, E.; Ricard, A. doi  openurl
  Title Optimization of breeding schemes for sport horses Type Journal Article
  Year 2008 Publication Livestock Science Abbreviated Journal  
  Volume 118 Issue 1-2 Pages 99-112  
  Keywords Breeding scheme; Horse; Jumping; Optimization; Genetic trend, Multistage selection  
  Abstract A selection scheme for jumping sport horses is modelled with four stages of selection for males and one stage for females. The selection objective included three traits: conformation and gaits (CG, weighted 20%), competition jumping (CJ, weighted 60%) and a third trait (TT, weighted 20%) such as sperm quality or orthopaedic status. The first selection stage is based on knowledge of the pedigree with the aim of selecting horses suitable for CG test (at 3 years old) and CJ test (at 5 years old). The second stage includes the horse's own performance with respect to CG and CJ with the aim of selecting horses suitable for the TT test. The third stage is the selection of a limited number of males who are allowed to reproduce. The fourth stage (at 12 years old) takes into account the results of the horse's progeny. Females are selected in one step, whatever the number of performances measured at 5 years old. The annual genetic response was 9.4% genetic standard deviation of the objective, 2.6% for CG, 9.0% for CJ and 1.5% for TT. Results showed that selection by progeny testing did not contribute much to genetic response (12% of progeny issued from proven sires), the female pathway represented 26% of genetic response, TT was difficult to improve when the genetic correlation was unfavourable (- 0.6% genetic standard deviation for - 0.20 genetic correlation), and should consequently be directed towards the use of molecular markers. When compared with a selection scheme involving a station test, genetic response was the same if the breeding values used for selection before entering the station test took into account the results of the relatives for CJ and CG. This revealed the importance of an extensive performance test (like for competition performance) when designing breeding schemes for sport horses.  
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  Series Editor Series Title Abbreviated Series Title  
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  ISSN 1871-1413 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4759  
Permanent link to this record
 
 
Author Christensen, J.W.; Zharkikh, T.; Ladewig, J. url  doi
openurl 
  Title Do horses generalise between objects during habituation? Type Journal Article
  Year 2008 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 114 Issue 3-4 Pages 509-520  
  Keywords Horse; Stimulus generalisation; Habituation; Fear  
  Abstract Habituation to frightening stimuli plays an important role in horse training. To investigate the extent to which horses generalise between different visual objects, 2-year-old stallions were habituated to feeding from a container placed inside a test arena and assigned as TEST (n=12) or REFERENCE horses (n=12). In Experiment 1, TEST horses were habituated to six objects (ball, barrel, board, box, cone, cylinder) presented in sequence in a balanced order. The objects were of similar size but different colour. Each object was placed 0.5m in front of the feed container, forcing the horses to pass the object to get to the food. TEST horses received as many 2min exposures to each object as required to meet a habituation criterion. We recorded behavioural reactions to the object, latency to feed, total eating time, and heart rate (HR) during all exposures. There was no significant decrease in initial responses towards a novel object with increasing object number, indicating that habituation was stimulus-specific. REFERENCE horses were exposed to the test arena without objects in the same period, and reactions of TEST and REFERENCE horses were subsequently compared in a fear-test (2m2.5m rubber mat placed under the feed container, forcing the horses to step on the mat to get food). There were no significant differences between the treatment groups, i.e. previous habituation of TEST horses to six visual objects did not reduce responses in a fear-test involving visual and tactile stimulation. Due to the lack of generalisation in Experiment 1, we designed a supplemental experiment (Experiment 2), in which REFERENCE horses were exposed to the same six objects except that object colour was kept constant. We found a significant reduction in response (behaviour and HR) with increasing object number, indicating that horses generalise between similarly coloured objects of varying shape. We conclude that a high degree of object similarity, e.g. identical colouring, appears to be crucial for object generalisation in horses.  
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  Series Editor Series Title Abbreviated Series Title  
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  ISSN 0168-1591 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4756  
Permanent link to this record
 
 
Author Whishaw, I.Q.; Sacrey, L.-A.R.; Gorny, B. url  doi
openurl 
  Title Hind limb stepping over obstacles in the horse guided by place-object memory Type Journal Article
  Year 2009 Publication Behavioural Brain Research Abbreviated Journal Behav. Brain. Res.  
  Volume 198 Issue 2 Pages 372-379  
  Keywords Horse place memory; Memory and hind limb stepping; Obstacle memory in horse; Obstacle stepping in horse; Place memory for stepping; Stepping in the horse; Working memory for stepping  
  Abstract An animal that has stepped over an obstacle with its forelimbs uses a memory of the obstacle to guide the hind limbs so that they also clear the obstacle, even in situations in which long pauses are introduced between forelimb and hind limb stepping. To further clarify the features of hind limb obstacle clearance memory, the present study examined hind limb obstacle clearance in the horse. A rider guided horses over obstacles and paused the horse over obstacles in tests that examined the relationship between forelimb and hind limb stepping, with the following results. First, the horses displayed memory for an obstacle as measured by hind limb lifting over the obstacle for durations lasting as long as 15 min. The response was not dependent upon ongoing visualization of the obstacle, as limb lifting was unaffected by visual occlusion with blinders, a blindfold, or by removing the obstacle during the pause. Second, previous experience of stepping over an obstacle led to pause-related hind limb lifting at the object's previous location even on trials for which there was no obstacle and so no preceding forelimb lifting. Third, whereas a horse would lift its hind limbs to clear two successively presented obstacles, replacing an obstacle before the horse after the forelimbs had cleared the obstacle prevented subsequent hind limb lifting at the obstacle's previous location. Taken together the results show that hind limb obstacle clearance is guided by a place-object memory. The results are discussed in relation to the differential sensory and memonic control of forelimb and hind limb stepping with the suggestion that place-object memory can guide hind stepping as well as overshadow working memory from front leg stepping.  
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  ISSN 0166-4328 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4754  
Permanent link to this record
 
 
Author Björk, N. url  openurl
  Title Is it possible to measure the welfare of the ridden horse? Type Manuscript
  Year 2008 Publication Abbreviated Journal  
  Volume Issue Pages  
  Keywords horse, welfare, training, learning, measure, assess  
  Abstract Since the time of domestication, humans have trained horses for the purpose of serving man. Different training methods have been developed throughout the centuries; some were developed with consideration for the horse's welfare, while others disregarded welfare to a great extent. Most present day training is based upon making the horse perform a desired behaviour through dominance and subordination. Although cooperative training techniques have gained popularity, everyday training lacks the application of learning theory or neglects the horse's learning capacities and their species' specific behaviour. Thus, the horse's welfare may be jeopardised.

The aim with this review is to consider methods that allow an objective assessment of the welfare of horses undergoing training. The review gives a brief insight into the history of horse training and handling. It proceeds with an overview of the horse"s learning abilities which is argued to be of paramount importance for effective training. The review then describes a few selected training techniques that are used today, based on negative and positive reinforcement, and discusses parameters from which it could be possible to assess the welfare of the ridden horse. The work concludes with suggestion for future		  
  Address  
  Corporate Author Thesis Bachelor's thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4749  
Permanent link to this record
 
 
Author Verrill, S.; McDonnell, S. url  doi
openurl 
  Title Equal Outcomes with and without Human-to-Horse Eye Contact When Catching Horses and Ponies in an Open Pasture Type Journal Article
  Year 2008 Publication Journal of Equine Veterinary Science Abbreviated Journal  
  Volume 28 Issue 5 Pages 309-312  
  Keywords Horse handling; Horse management; Eye contact; Interspecies interactions; Equine behavior  
  Abstract Each of 104 horses and ponies was approached for catching at pasture by the same human handler in a standard manner, either maintaining human-to-animal eye contact (EC+; n = 51) or avoiding eye contact (EC-; n = 53). A subset of 74 of these subjects were reevaluated 3 weeks later under similar standard conditions except with the eye contact condition opposite to that used in the first round. Nonparametric statistical methods were used to evaluate between subjects (round 1, n = 104) and within subjects (rounds 1 and 2, n = 74) comparisons of successful or unsuccessful catching outcome with EC+ and EC-. Catching outcomes were similar with eye contact condition. Although this study represents a single handler at one study site, results suggest that human-to-horse eye contact may not be an important influence on catching pastured horses. Certainly, further work is needed to better understand the role of eye contact in horse handling.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0737-0806 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4711  
Permanent link to this record
 
 
Author Lansade, L.; Bouissou, M.-F. url  doi
openurl 
  Title Reactivity to humans: A temperament trait of horses which is stable across time and situations Type Journal Article
  Year 2008 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 114 Issue 3-4 Pages 492-508  
  Keywords Horse; Equus caballus; Reactivity to humans; Temperament trait; Personality; Behavioural consistency  
  Abstract The aim of this study was to test the existence of a “reactivity-to-humans” trait which is stable over time and across situations. For this purpose, four test situations were repeated on the same animals at 8 months, 1.5 years (N = 110 horses) and 2.5 years of age (N = 54 horses). These situations involved a “familiar passive human test” during which a familiar person stayed motionless in the test pen, a “familiar active human test” and an “unfamiliar active human test” during which a familiar or unfamiliar person attempted to touch the horse, and a “halter-fitting and heart-rate measurement test”, during which the horse was haltered and equipped with a surcingle and then had its heart-rate measured. Many behavioural variables were recorded during the tests, and correlations between ages and variables measured during other situations were investigated. When correlations were found, we concluded that the variables corresponded to temperament traits which are stable over time and across situations. The horses tested were divided into four groups according to breed (Anglo-Arab and Welsh) and year of birth (2001 and 2002). Data for each group were analysed separately with Spearman rank correlations. Regarding stability over time, there was a strong correlation between ages in the frequency of licking/nibbling the passive human, the time taken by a familiar or unfamiliar person to touch the horse, and the time taken to fit a halter (e.g. the time taken to touch the horse's muzzle was significantly correlated between ages in 7 out of the 8 possible cases: 0.40 <= R <= 0.67). Regarding stability across situations, results indicate that the more frequently a horse sniffed, licked or nibbled a passive human, the easier it was for either a familiar or unfamiliar active human to touch and halter it. Moreover, a horse which could be touched easily by a familiar human could also be touched and haltered easily by an unfamiliar human. These behaviours, which are linked directly to humans, therefore indicate good stability over time and across situations. In conclusion, this stability suggests the existence of a “reactivity-to-humans” temperament trait, whether the human is passive or active, familiar or unfamiliar. When the animals do not have frequent contact with humans, this reactivity is stable over time and can be measured as early as 8 months of age using the behavioural variables cited above.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0168-1591 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4710  
Permanent link to this record
 
 
Author Cameron, E.Z.; Linklater, W.L.; Stafford, K.J.; Minot, E.O. url  doi
openurl 
  Title Maternal investment results in better foal condition through increased play behaviour in horses Type Journal Article
  Year 2008 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.  
  Volume 76 Issue 5 Pages 1511-1518  
  Keywords Equus caballus; feral horse; maternal investment; play  
  Abstract Play behaviour is widespread in mammals, but benefits to play have been difficult to demonstrate. Physical training is one of the many proposed hypotheses, suggesting that males and females should play differently, that increased maternal investment should lead to increases in play, and that increases in play should result in physical advantages. In a population of feral horses, Equus caballus, males and females did not differ in their play behaviour except that males initiated more of their play bouts than females. Maternal condition influenced play behaviour only in males, with sons of mothers in good condition playing more. However, when we controlled for maternal effects by comparing a son and a daughter of the same mother, daughters played more when their mother was in poor condition and sons played more when their mother was in good condition. Mothers of foals that played more lost more condition. Therefore, the difference in play behaviour could not be explained by offspring sex or maternal condition alone, but play behaviour mirrored variation in maternal investment. In addition, those individuals that played more survived better and had better body condition as yearlings despite weaning earlier. Since increased activity has been linked to enhanced musculoskeletal development in domestic horses, we suggest that play provides a link between increased maternal investment, increased body condition and future reproductive success in feral horses, and probably in other species.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0003-3472 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4709  
Permanent link to this record
 
 
Author King, S.R.B. url  doi
openurl 
  Title Home range and habitat use of free-ranging Przewalski horses at Hustai National Park, Mongolia Type Journal Article
  Year 2002 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 78 Issue 2-4 Pages 103-113  
  Keywords Przewalski horse; Equus przewalskii; Takhi; Home range; Re-introduction; Habitat use  
  Abstract Przewalski horses (Equus ferus przewalskii), also known as takhi, were first re-introduced to the wild in Hustai National Park, Mongolia, in 1994. Since then the number of free harems increased to a maximum of seven; there are currently six (October 2000). The size of the home range of each of the harems changed among years and among seasons. The horses tended to settle in a home range close to where they were released although they explored the surrounding area. The use of the habitat within each home range changed through the day, with the horses grazing in the valleys during the morning and evening, and moving to higher places to stand rest and use as a refuge from heat and flies during the middle of the day. Range establishment and area, as well as habitat use are discussed.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0168-1591 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4682  
Permanent link to this record
 
 
Author Gatta, D.; Baragli, P.; Ferrarini, N.; Ciattini, F.; Sighieri, C.; Colombani, B. openurl 
  Title Fitness evaluation in endurance horses by standardised exercise test on treadmill Type Journal Article
  Year 1998 Publication Ippologia Abbreviated Journal Valutazione dello stato di allenamento del cavallo da endurance mediante test standardizzato su trea  
  Volume 9 Issue 4 Pages 57-69  
  Keywords Endurance; Horses; Treadmill  
  Abstract Endurance horses, like marathon runners, undertake sub-maximal, longterm physical exercises and in both cases thermoregulation plays a critical role. A standardised sub-maximal treadmill test was used to assess fitness and training state in endurance horses. Literature reports that trained human athletes dissipate head load better than not trained ones; in this study parameters related to thermoregulation were recorded together with other haematologic and metabolic ones. Six endurance horses, divided into two groups according to their training state, performed two submaximal tests at a distance of one week. Blood samples were collected at rest, during exercise, at the end and during recovery (10, 30 and 60 minutes after work). Heart rate (HR), respiratory rate (RR), rectal (RT) and skin (ST) temperatures were monitored through the exercise (and recovery for HR). HR resulted significantly lower (P<0.01) in trained horses than in untrained ones, especially during recovery. Also rectal and skin temperatures resulted significantly lower (P<0.05) in trained horses and ST increased more gradually during exercise. The values of haematocrit and haemoglobin resulted significantly lower (P<0.05) at rest, during exercise and recovery samples. In this study, the parameters related to thermoregulation, in accordance with data from human athletes, resulted significantly different in trained and untrained horses, while other parameters didn't vary significantly between the two groups. Thermoregulatory responses to exercise-generated heat load in a standardised test, studied by simple parameters, can thus offer useful information about fitness and training state of endurance horses.  
  Address Dipto. Anat., Biochim. Fisiol. Vet., Univ. degli Studi di Pisa  
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  Notes Cited By (since 1996): 2; Export Date: 13 November 2008; Source: Scopus Approved no  
  Call Number Equine Behaviour @ team @ Serial 4655  
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