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	Author	Krueger, K.; Esch, L.; Byrne, R.
	Title	Need or opportunity? A study of innovations in equids			Type	Journal Article
	Year	2021	Publication	Plos One	Abbreviated Journal	Plos One
	Volume	16	Issue	9	Pages	e0257730
	Keywords
	Abstract	Debate persists over whether animals develop innovative solutions primarily in response to needs or conversely whether they innovate more when basic needs are covered and opportunity to develop novel behaviour is offered. We sourced 746 cases of “unusual” behaviour in equids by contacting equid owners and caretakers directly and via a website (https://innovative-behaviour.org), and by searching the internet platforms YouTube and Facebook for videos. The study investigated whether differences in need or opportunity for innovation were reflected in the numbers of different types of innovations and in the frequencies of repeating a once-innovative behaviour (i) with respect to the equids' sex, age, and breed type, (ii) across behavioural categories, and whether (iii) they were affected by the equids' management (single vs group housing, access to roughage feed, access to pasture, and social contact). We found that the numbers of different types of innovation and the frequency of displaying specific innovations were not affected by individual characteristics (sex, age, breed or equid species). Few types of innovation in escape and foraging contexts were observed, whilst the comfort, play, and social contexts elicited the greatest variety of innovations. We also found higher numbers of different types of innovations in horses kept in groups rather than in individual housing, and with unlimited rather than with restricted access to pasture and roughage. Equids in permanent social contact performed high rates of once-innovative behaviour. We suggest that equids produce goal-directed innovations and repeat the behaviour at high frequency in response to urgent needs for food and free movement or when kept in conditions with social conflict. However, equids devise the greatest variety of innovations when opportunity to play and to develop comfort behaviour arises and when kept in good conditions.
	Address
	Corporate Author				Thesis
	Publisher	Public Library of Science	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6653
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	Author	Krueger, K.; Gruentjens, T.; Hempel, E.
	Title	Wolf contact in horses at permanent pasture in Germany			Type	Journal Article
	Year	2023	Publication	Plos One	Abbreviated Journal	Plos One
	Volume	18	Issue	8	Pages	e0289767
	Keywords
	Abstract	Wolves returned to Germany in 2000, leading to fear in German horse owners that their horses could be in danger of wolf attacks or panic-like escapes from pastures when sighting wolves. However, reports from southern European countries indicate that wolf predation on horses diminishes with increasing presence of wildlife. Therefore, we conducted a long-term, filed observation between January 2015 and July 2022 on 13 non breeding riding horses, mares and geldings, kept permanently on two pastures within the range of wildlife and a stable wolf pack with annual offspring. Wildlife cameras at the fences of the pastures made 984 times recordings of wolves and 3151 times recordings of wildlife in and around the pastures. Between 1 January 2022 and 23 March 2022 we observed two stable horse groups. Pasture 1 was grazed by five horses of mixed breed, four mares and one gelding, with the median age of 8 years (min. = 6y, max. = 29y). Pasture 2 was grazed by eight heavy warmbloods and draught horses, three mares and five geldings, with the median age of 16 years (min. = 13y, max. = 22y). During this period no wolf was recorded at pasture 2, but wild boar several times, whereas at pasture 1, wolves were recorded 89 times, and for the wildlife mostly hare. Wolves may have avoided pasture 2 because of the presence of wild boar or because the large group of older, heavy breed horses may have formed a stable, protective group. The latter needs to be confirmed in a follow-up field observation, which records anti-predator behavior and welfare indicators in horses. In conclusion, wolves did not attack the mature horses on pastures with plenty of wildlife and the horses did not respond to the presence of wolves with visible signs of reduced welfare or panic. This indicates that wolves may prefer to prey on easily accessible wildlife around and at horse pastures and that Central European horses become accustom to the presence of non-hunting wolves.
	Address
	Corporate Author				Thesis
	Publisher	Public Library of Science	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6708
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	Author	Virányi, Zs.; Range, F.; Huber, L.
	Title	Attentiveness toward others and social learning in domestic dogs.			Type	Book Chapter
	Year	2008	Publication	Learning from Animals?: Examining the Nature of Human Uniqueness	Abbreviated Journal
	Volume		Issue		Pages	141-154
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Psychology Press	Place of Publication	New York, NY	Editor	Röska-hardy,L.S.. ;Neumann-held, E.
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN	978-1-84169-707-9	Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4974
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	Author	Croft, D. P.; James, R..; Krause, J. (eds)
	Title	Exploring Animal Social Networks			Type	Book Whole
	Year	2008	Publication		Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Princton University Press	Place of Publication	Princton	Editor	Croft, D. P.; James, R..; Krause, J.
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN	9780691127521	Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5139
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	Author	Wynne C. D. L.
	Title	Animal Cognition: The Mental Lives of Animals			Type	Book Whole
	Year	2001	Publication		Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract	Covering a wide range of key topics, from reasoning and communication to sensation and complex problem-solving, this engagingly-written text presents a comprehensive survey of contemporary research on animal cognition. Written for anyone with an interest in animal cognition, but without a background in animal behaviour, it endeavours to explain what makes animals tick. With numerous illustrations and including exciting recent studies from many little-studied species (such as the weakly electric African fish), this text is ideal for psychology students who are interested in how much of our human cognition is shared by other species, for students of biology who want to know how complex animal behaviour can get, and for all those with an interest in the animal mind.
	Address
	Corporate Author				Thesis
	Publisher	Palgrave	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN	9780333923955	Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6157
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	Author	Van Horik, J.; Clayton, N.; Emery, N.
	Title	Convergent evolution of cognition in Corvids, Apes and other animals			Type	Book Chapter
	Year	2012	Publication	Oxford Handbook of Comparative Evolutionary Psychology	Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Oxford University Press	Place of Publication	New York	Editor	Vonk, J.; Shackelford, T.
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Van Horik2012			Serial	6284
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	Author	Van Horik, J.; Clayton, N.; Emery, N.
	Title	Oxford Handbook of Comparative Evolutionary Psychology			Type	Book Whole
	Year		Publication		Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Oxford University Press	Place of Publication	New York	Editor	Vonk, J.; Shackelford, T.
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6403
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	Author	Reader, S.M.; Laland, K.N.
	Title	Animal Innovation			Type	Book Whole
	Year	2009	Publication		Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Oxford University Press	Place of Publication	Oxford	Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN	978-0-19-852622	Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6381
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	Author	Reader, S. M.; Laland, K.N.
	Title	Animal Innovation			Type	Book Whole
	Year	2003	Publication		Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Oxford University Press	Place of Publication	Oxford	Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6531
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	Author	Sol, D.
	Title	Behavioural flexibility: a neglected issue in the ecological and evolutionary literature			Type	Book Chapter
	Year	2003	Publication	Animal innovation.	Abbreviated Journal
	Volume		Issue		Pages	63-82
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Oxford University Press	Place of Publication	Oxford	Editor	S. M. Reader and K. N. Laland
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6532
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